Employment

		2015-present	Professor of Vision Science, Newcastle University, UK.
		2013-2015	Reader in Vision Science, Newcastle University, UK.
		2005-2013	Royal Society University Research Fellow, Newcastle University, UK.
		2001-2005	Post-doctoral Fellow, Laboratory of Sensorimotor Research, National Eye Institute, National Institutes of Health, MD
		1997-2001	Wellcome Trust Mathematical Biology Training Fellow, University Laboratory of Physiology, Oxford University
		2000-2001	College lecturer in physics at Corpus Christi College, Oxford
		1999-2000	College lecturer in physics at Christ Church, Oxford
		1994-1997	UK Particle Physics and Astrophysics Research Council Studentship
		1991-1994	UK Atomic Energy Authority Studentship

Qualifications

		2006-2009	Graduate Diploma in Psychology (with distinction), Newcastle University.
		1998-1999	M.Sc. in Neuroscience (with distinction), University of Oxford, Merton College.
		1994-1997	D. Phil. in Theoretical Physics, University of Oxford, Merton College. Thesis entitled “The stability of model disk galaxies”
		1991-1994	B. A. First Class Honours in Physics, University of Oxford, University College.
		1996-2001	Various courses in biology, maths, German, The Open University.
		A-levels:	Latin (Grade A, 1990); Ancient Greek (grade A, 1990); Physics (grade A, 1991); Mathematics (grade A, 1991); Further Mathematics (grade A, 1991); Russian (grade A, 2008)